Table 2 Results of two GLMMs testing the influence of different landscape attributes (slope, presence of canopy gaps, elevation and FTs density) on the probability occurrence of a route segment used at least twice and four times within a certain quadrat. Group ID (N = 5) was included as random effect in the model. We compared the full model to a corresponding null model (with only random and control variables) using likelihood-ratio tests. All p-values < 0.05 are shown in bold for clarity